Graduate Student Cohorts


Interests and Research Projects


Get to know the past and present students of the Primate Behavior MS Graduate Program. Read theses published by former students and learn about the variety of research projects completed or in progress by CWU students and their faculty advisors.
  • 2014 Cohort

    Krista (Kris) Banda, BA Anthropology & Psychology, Emory University
    Faculty advisor: Dr. Gabrielle Stryker
    Research interests: She arrived at her interest in primate parasite ecology through a circuitous path that started by working in a neuroscience lab to being a primate intern to working in a Dian Fossey Gorilla Fund International lab. Her research in Peru focused on the intestinal parasites of saddleback (Saguinus fuscicollis) and emperor tamarins (S. imperator).
    Thesis: The Seasonality and Parasite and Prevalence of the Weddelli’s Saddleback Tamarin (Leontocebus Weddelli) 

    Allison Farley, BA Environmental Science, minor in Biology, Sonoma State University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: She previously interned in the Bay Area monitoring populations of endemic endangered species. While at Central she volunteered at Chimpanzee Sanctuary Northwest.
    Thesis: Comparison of chimpanzee (Pan troglodytes) behavior on tour and non-tour days at Chimpanzee Sanctuary Northwest / 2016 

    Ruth Linsky
    Faculty advisors: Dr. Joseph Lorenz & Dr. Steve Wagner
    Thesis: A Novel SNP Genotyping Technique to Determine Orangutan Relatedness, Kinship, and Genetic Diversity at Camp Leakey in Tanjunf Puting, Central Kalimantan / 2019 

    Brianna Schnepel, BA Anthropology, San Diego State University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Brianna has previously worked with white-faced capuchin monkeys (Cebus capucinus) and mantled howler monkeys (Alouatta palliata) while attending a field school in Costa Rica. As an undergraduate, she also looked at interactions between bonobos (Pan paniscus) and human visitors at the San Diego Zoo while assisting a graduate student from San Diego State University. Her thesis research focused on the effects that human food had on the social interactions and behavior of Tibetan macaques (Macaca thibetana) living at a tourist site at Mt. Huangshan, China. Brianna's fieldwork was funded by a grant from the National Science Foundation. Her faculty advisor was Dr. Lori Sheeran.
    Thesis: Provisioning and its effects on the social interactions of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China / 2016 

    Holly Soubiea, BA Psychology, minor in Biology, Holy Names University
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Holly has previously been an Animal Care Intern at the Oakland Zoo, where she worked with chimpanzees (Pan troglodytes), squirrel monkeys (Saimiri sciureus), hamadryas baboons (Papio hamadryas), and gibbons (Hylobates lar). She was a volunteer at Chimpanzee Sanctuary Northwest while at Central. Her thesis research focused on chimpanzees' use of their enclosures at Fauna Foundation.
    Thesis: Chimpanzee (Pan troglodytes) Space Use in a Sanctuary Setting / 2017 

    Emily Veitia, BA Sociology/Anthropology, minor in Geography, Florida International University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: She previously volunteered at Chimpanzee Sanctuary Northwest and currently volunteers at the Mimulus Guttatus lab with Dr. Alison Scoville. Emily's thesis research focused on cross-species interactions in a mixed-species ape exhibit at the Oregon Zoo.
    Thesis: The effect of enrichment types in a mixed-species enclosure of Sumatran (Pongo abelii) & Bornean Orangutans (Pongo pygmaeus) and Northern White-Cheeked Gibbons (Nomascus leucogenys) / 2017 

    Kaitlin Wright, BA Biological Anthropology, minor in Biology, University of San Diego
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Kaitlin has worked with Orangutan Foundation International in Kalimantan, Indonesia and the Pacific Primate Sanctuary on Maui, Hawaii. Her thesis research was conducted at the Valley of the Wild Monkey in Mt. Huangshan, China and focused on the rate of play signals in Tibetan macaques (Macaca thibetana) during juvenile play bouts. Kaitlin's fieldwork was funded by a grant from the National Science Foundation.
    Thesis: Play in Immature Tibetan Macaques (Macaca thibetana): Location, Use of Play Signals, and Play Bout Termination at Mt. Huangshan, China / 2016
  • 2013 Cohort

    Gregory Fratellone, BS Biology, minor in Psychology, Northeastern University
    Faculty advisor: Dr. Lixing Sun
    Research interests: Greg previously conducted research on western lowland gorillas (Gorilla gorilla gorilla) at the Franklin Park Zoo, focusing on infant development and personality, and has volunteered in Belize at the Belize Zoo and Wildtracks rehabilitation center with Geoffroy's spider monkeys (Ateles geoffroyi) and Yucatan black howler monkeys (Alouatta pigra). His thesis research was conducted at the Valley of the wild Monkeys in Mt. Huangshan, China, examining the collective movements of Tibetan macaques (Macaca thibetana) and analyzing collective decision-making, leader/follower dynamics and social networks.
    Thesis: Female Social Connectivity Through the Leadership and Movement Progression of Tibetan Macaques at Mt. Huangshan, China / 2015 

    Melanie Bell, BA Anthropology, BS Primate Behavior & Ecology, Central Washington University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Melanie is a former intern at the Chimpanzee and Human Communication Institute (CHCI). Her thesis research was performed at the Gibbon Conservation Center in southern California looking at the non-vocal communication of Javan Gibbons (Hylobates moloch).
    Thesis: Javan Gibbon (Hylobates moloch) Non-Vocal Social Communication and Gesture Use with Conspecifics / 2015 

    Whitney Emge, BA Psychology, University of Puget Sound
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Whitney was an apprentice at the Chimpanzee and Human Communication institute (CHCI) and intern at Chimpanzee Sanctuary Northwest. Her thesis research was conducted at Fauna Foundation in Carignan, Quebec, looking at the effects of operant training sessions and unstructured interactions between chimpanzees (Pan troglodytes) and their caregivers.
    Thesis: A Comparison of Chimpanzee (Pan troglodytes) Response to Caregiver Use of Positive Reinforcement Training (PRT) and Species-Specific Behaviors (SSB) / 2015 

    Katherine McDonald, BA Psychology & Biological Sciences, Wellesley College
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Katie previously worked with tufted capuchin monkeys (Cebus apella) at Yale University, examining helping behavior between conspecifics. She was an apprentice at the Chimpanzee and Human Communication Institute (CHCI) and intern at Chimpanzee Sanctuary Northwest. Katie is interested in primate cognition, primate prosocial behavior, and captive primate welfare.
    Thesis: Captive Chimpanzee (Pan troglodytes) Nighttime Enrichment Use and Preferences / 2016 

    Alexandra Casti, BFA Painting, minor in Art History, Pratt Institute
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Sandra was an apprentice at the Chimpanzee and Human Communication Institute (CHCI) and intern/volunteer caregiver at Chimpanzee Sanctuary Northwest. She is passionate about personhood rights for nonhuman animals and interested in cognitive development, creative expression, and language evolution. She analyzed a series of chimpanzee (Pan troglodytes) drawings from the CHCI archive for her thesis.
    Thesis: Reaction to Stimulus Figures in Chimpanzee (Pan troglodytes) Drawings / 2016 

    Benjamin Gombash, BS Wildlife Sciences & Anthropology, The Ohio State University
    Faculty advisor: Dr. Joseph Lorenz
    Research interests: Ben is interested in primate conservation and vervet monkeys (Chlorocebus aethiops). His research concerned Hepatocystis and how genes might confer resistance to it.
    Thesis: Analysis of FY Promoter and Hepatocystis Load in South African Vervet monkeys (Chlorocebus aethiops) / 2016 

    Anne Salow
    Faculty Advisor: Dr. Steve Wagner
    Thesis: Sexual Behavior of Immature Tibetan Macaques (Macaca thibetana) / 2015 

  • 2012 Cohort

    Amanda Carner
    Faculty Advisor: Dr. Mary Lee Jensvold
    Thesis: Gesture Sequence in Captive Orangutans (Pongo pygmaeus) / 2014

    Tony Carr
    Faculty Advisor: Dr. Marte Fallshore
    Thesis: Empathy: A Possible Link Between Charismatic Megafauna and Environmental Concern / 2014

    Ricardo Fernandes, BS Animal & Veterinary Science, minor in Biology & Theatre Arts, University of Rhode Island
    Faculty advisor: Dr. Joseph Lorenz
    Research interests: Ricardo is a licensed veterinary technician and specializes in emergency and critical care medicine. His areas of interest include primate communication, primate physiology of locomotion, infant/mother interactions and parental care. In addition, Ricardo is interested in the welfare of captive primates. He analyzed fecal samples to look at population genetics of white faced saki monkeys (Pithecia pithecia) in South America.
    Thesis: Sex differences in glycosylated hemoglobin in Mauritian origin long-tailed macaques (Macaca fascicularis) / 2019 

    Erika Jackson Price,  BA Neuroscience, minor in Jazz Music, University of Southern California
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Erika's primary interests are the behaviors of free-ranging primates, psychology, and music. She recorded, cataloged, and analyzed the vocalizations of Tibetan macaques at Mt. Huangshan, China.
    Thesis: Vocalizations of Tibetan Macaques (Macaca thibetana) at Mt. Huangshan, China / 2014

    Savannah Schulze
    Faculty advisor: Dr. Mary Lee Jensvold
    Thesis: The Directed Scratch Gesture: A Referential Gesture in Chimpanzees / 2014
    Jensvold ML; Wilding L; Schulze SM. 2014. Signs of communication in chimpanzees. G. Witzany (Ed.) Biocommunication of Animals (pp. 7-19). Dordrecht: Springer.

    RyAnn Stafford
    Faculty Advisor: Dr. Mary Lee Jensvold

    Asa Staven, BS Fish & Wildlife Management, minor in Entomology, Montana State University
    Thesis advisor: Dr. Steven Wagner
    Research interests: Asa had a Teaching Assistantship from the Biology Department and studied the behaviors of juvenile Tibetan macaques (Macaca thibetana).
    Thesis: Interactions Between Tourists and Juvenile Tibetan Macaques (Macaca thibetana) at Mt. Huangshan, China / 2014

    Kaeley Sullins
    Faculty Advisor: Dr. Mary Lee Jensvold
    Thesis: Effect of Group Size on the Activity Budget of Two Captive Chimpanzees (Pan troglodytes) / 2019 

    Natasha Zientek, BA Anthropology, The Ohio State University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Natasha studied community-based primate conservation focused on Tibetan macaques at Mt. Huangshan, China.
    Thesis: Impacts of Behavioral Education on Actions and Conservation Attitudes of Tourists at the Valley of the Wild Monkeys, Mt. Huangshan, China / 2014

  • 2011 Cohort

    Alexander DuVall-Lash, BS Evolutionary Anthropology, University of Michigan
    Faculty advisor: Dr. Lori Sheeran
    Alexander studied the impact of tourism on stress-indicating behaviors in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
    Thesis: Effect on human noise on stress-indicating behaviors of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China / 2013 

    Jennifer Humphreys, BA Anthropology emphasis on Biological Anthropology, Humboldt State University
    Faculty advisor: Dr. Joseph Lorenz
    Jennifer studied conservation population genetics, comparative skeletal anatomy, behavioral ecology, and ethnoprimatology.
    Thesis: DNA from Chewed Seeds: A Non-Invasive Collection Strategy for Saguinus midas / 2014

    Susie Keenan, BS Psychology, Rochester Institute of Technology
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Susie worked at the Chimpanzee Human Communication Institute and is interested in gestural communication and American Sign Language.

    Meg Mas, BS in Primate Behavior, BA Psychology, Central Washington University
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Meg worked at the Chimpanzee Human Communication Institute and is interested in imaginary play in nonhuman primates.

    Alex Pritchard, BS Biology, University of New South Wales
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Alex is primarily interested in the social behaviors and social interactions of all primates, but also in bolstering their protection, preservation, and recognition.
    Thesis: Discriminating Behaviors to Measure Personality Traits in Free-Ranging Tibetan Macaques (Macaca thibetana) / 2013

    Leah Usui, BS Evolution & Ecology and Anthropology, The Ohio State University
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Leah is interested in the effect of site management styles and ecotourism impact on Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
    Thesis: Behaviors of park rangers and their effects on Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China / 2013 
    Usui R, Sheeran Lk, Li JH, Sun L, Wang X, Pritchard AJ, DuVall-Lash AS, Wagner RS. 2014. Park rangers' behaviors and their effects on tourists and Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. Animals 4:546-561, doi: 10.3390/ani4030546.

    Lisa Wilding, BA Ecology & Evolutionary Biology, University of Colorado at Boulder
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Lisa worked at the Chimpanzee Human Communication Institute and is interested in gorilla communication.
    Thesis: Gorilla Life-Stage Comparison of Head Orientation / 2017 
    Jensvold Ml; Wilding L; Schulze SM. 2014. Signs of communication in chimpanzees. G. Witzany (Ed.), Biocommunication of Animals (pp. 7-19). Dordrecht: Springer.

  • 2010 Cohort

    Briana Bauer, BA Spanish, University of Notre Dame
    Faculty advisor: Dr. Lori Sheeran
    Research interest: Briana studied bridging behaviors and adult male-adult female affiliative relationships in Tibetan macaques (Macaca thibetana) at the Valley of the Wild Monkeys in Mt. Huangshan, China. During her time at CWU, she finished internships at the Woodland Park, Toledo, and Roger Williams Park zoos.
    Thesis: Male Tibetan Macaque (Macaca thibetana) Choice of Infant Bridging and Female Affiliation Partners / 2012
    Bauer B; Sheeran LK; Matheson MD; Li JH; Wagner RS. 2014. Choice of infant bridging and female affiliation partners by male Tibetan macaques (Macaca thibetana). Zoological Research 35(3):222-230.

    Sofia Bernstein, BA Psychology emphasis in Applied Behavioral Analysis, University of North Texas.
    Faculty advisors: Dr. Lori Sheeran & Dr. Steve Wagner
    Research interest: Sofi studied sexual coercion and mate choice in Tibetan macaques (Macaca thibetana) at the Valley of the Wild Monkeys in Mt. Huangshan, China.
    Thesis: Sexual Coercion and Mate Choice Tactics in Tibetan Macaques (Macaca thibetana) at Mt. Huangshan, China / 2012

    Laura Burwell, BS Wildlife and Wildlands Conservation, Brigham Young University
    Faculty advisor: Dr. Lori Sheeran
    Research interest: Laura studied competition for provisioned foods in a group of Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
    Thesis: The Effectiveness of Scattering Corn as a Management Technique for Macaca thibetana at Mt. Huangshan, China / 2013

    Amanda Davis, BS Anthropology & Liberal Studies, Portland State University
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interest: Amanda studied the effects of conversational partner familiarity in deaf humans.
    Thesis: Effects of Conversational Partner Familiarity in Deaf Humans (Homo sapiens) / 2012

    Erica Dunayer, BS Animal Behavior, Bucknell University
    Faculty advisor: Dr. Megan Matheson
    Research interest: Erica studied how tourism affects the grooming for tolerance trade in Tibetan macaques (Macaca thibetana) from a biological markets perspective at Mt. Huangshan, China. 
    Thesis: Tourism as a Market Force for Grooming Interchange and Reciprocity in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China / 2012

    Laura Heggs, BA Psychology, minor in Anthropology/Sociology, Knox College
    Faculty advisor: Dr. Megan Matheson
    Research interest: Laura studied the effect a novel outdoor environment had on captive chimpanzees at Chimpanzee Sanctuary Northwest in Cle Elum, Washington. 
    Thesis: The Effect of a Novel Outdoor Environment on the Behavior of Chimpanzees (Pan troglodytes) in a Sanctuary Setting / 2012

    Allie Kibler-Campbell, BA Psychology, minor in English, College of Wooster
    Faculty advisor: Dr. Lori Sheeran
    Research interest: Allie studied the ability to infer causality through various tool-use tasks (including stick tools and rock tools) in a colony of captive capuchin monkeys (Cebus apella) in Wooster, Ohio.
    Thesis: Causal Inference Examined in Cebus apella through a Hammer-and-Anvil Task / 2012

    Austin Leeds, BS Biology, BS Primate Behavior & Ecology, Central Washington University
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interest: Austin studied language use of the cross-fostered chimpanzees at the Chimpanzee Human Communication Institute.
    Thesis: The Communicative Functions of Five Signing Chimpanzees (Pan troglodytes) / 2012
    Leeds CA; Jensvold ML. 2013. The communicative functions of five signing chimpanzees (Pan troglodytes). Pragmatics & Cognition, 21(1):224-247

    Julie Reveles, BS Anthropology and Primate Behavior & Ecology, Central Washington University
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interest: Julie studied visitor effects on captive chimpanzees at the Chimpanzee Human Communication Institute.
    Thesis: Assessing the Reliability of a Written Data Collection Method: Sign Logs / 2014

    Noah Simons, BA Biological Anthropology, University of Iowa
    Faculty advisors: Dr. Steve Wagner & Dr. Joseph Lorenz
    Research interest: Noah studied levels of genetic diversity between captive and wild populations of western lowland gorillas.
    Thesis: Genetic diversity in captive-born gorillas (Gorilla gorilla gorilla) / 2012
    Simons ND; Wagner RS; Lorenz JG. 2013. Genetic diversity of North American captive-born gorillas (Gorilla gorilla gorilla). Ecology and Evolution, 3(1):80-88.

    Hilaree Sorenson, BA International Studies, Pepperdine University
    Faculty advisor: Dr. Mary Lee Jensvold
    Hilaree designed and implemented an environmental enrichment protocol for captive baboons at the Centre for Animal Rehabilitation & Education (C.A.R.E.) in Phalaborwa, South Africa.

  • 2009 Cohort

    Daniella Bismanovsky, BS Experimental Psychology, Saint Mary's College of California
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interest: Daniella's thesis examined the effect of visitors' behaviors on the chimpanzees' behaviors at the Oakland Zoo in California.
    Thesis: Chimpanzee (Pan troglodytes) Responses to Visitors Using Chimpanzee-Friendly Behaviors / 2012 

    Tracy Campion, BA Social Sciences, minor in Anthropology, University of Washington
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interest: Tracy examined gesture modality in relation to the attentional state in chimpanzees at Gombe in Tanzania, Africa. 
    Thesis: Use of Gesture Sequences in Free-Living Chimpanzee (Pan troglodytes schweinfurthii) Play in Gombe National Park, Tanzania / 2012
    Campion TL; Jensvold ML; Larsen G. 2011. Use of gesture sequences in free-living chimpanzees (Pan troglodytes schweinfurthii) in Gombe National Park, Tanzania. American Journal of Primatology,73(S1): 97

    Grace Enlow, BA Anthropology & Study of Religion, University of California, San Diego
    Faculty advisor: Dr. Lori Sheeran
    Research interest: Grace studied vocalizations and pair-bond strength in wild, white-bearded gibbons (Hylobates agilis albiarbis) in Central Kalimantan, Borneo.
    Thesis: Singing, Behavioral Synchrony, and Proximity in Bornean Agile Gibbons (Hylobates albibarbis) / 2011

    Rebecca Hendershott, BS Ecology & Evolutionary Biology, University of California, Santa Cruz
    Faculty advisor: Dr. Megan Matheson
    Research interest: Rebecca studied the sociosexual behaviors of Tibetan macaques (Macaca thibetana) at the Valley of the Wild Monkeys in Mt. Huangshan, China.
    Thesis: Socio-sexual Behavior in Tibetan Macaques (Macaca thibetana) / 2011

    Julie Wescliff, BA Psychology, University of Louisville
    Faculty advisor: Dr. Megan Matheson
    Research interest: Julie studied the impact of proximity on behavior in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China.
    Thesis: How Close is Too Close? Spatial Proximity Dynamics of Macaca thibetana / 2012

    Sandra Winters, BS Psychology & Anthropology, University of Maryland at College Park
    Faculty advisor: Dr. Megan Matheson
    Research interest: Sandra studied recruitment behavior in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China, for evidence of triadic awareness.
    Thesis: Social Recruitment in Tibetan Macaques (Macaca thibetana) at Mt. Huangshan, China / 2011

  • 2008 Cohort

    John Rinear, BA Anthropology, BS Primate Behavior & Ecology, Central Washington University.
    Faculty advisor: Dr. Lori Sheeran
    Research interests: John studied the sleeping site preferences of wild white-bearded gibbons (Hylobates agilis albiarbis).
    Thesis: Sleeping Tree Preference in Bornean Agile Gibbons (Hylobates albibarbis) / 2014
    Cheyne SM, Hoing A, Rinear J, Sheeran LK. 2012. Sleeping site selection by agile gibbons: The influence of tree stability, fruit availability and predation risk. Folia Primatologica, 83:299-311, DOI: 10.1159/000342145.

    Sydney Self, BS Anthropology, BS Primate Behavior & Ecology, Central Washington University.
    Faculty advisor: Dr. Lori Sheeran
    Research interests: Sydney studied infant-directed aggression in Tibetan macaques (Macaca thibetana) at Valley of the Wild Monkeys in Mt. Huangshan, China.
    Thesis: Tourist Density and Infant-Directed Aggression in Macaca thibetana at Mt. Huangshan, China / 2010
    Self S; Sheeran LK; Matheson MD; Li JH; Pelton O; Harding S; Wagner RS. 2013. Tourism and infant-directed aggression in Tibetan macaques (Macaca thibetana) at Mt. Huangshan, China. Anthrozoös, 26(3):435-444.

    Lindsay Zager 
    Faculty advisor: Dr. Mary Lee Jensvold
    Research interests: Lindsay studied visitor effects in zoo-living chimpanzees.
    Thesis: An Experiment in Zoo Visitor Education: Encouraging Friendly Chimpanzee (Pan troglodytes) Behaviors / 2011
    Jensvold ML; Zager L; Bismanovsky D. 2013. Promoting nonhuman animal welfare: interactions with caregivers and zoo visitors. Journal of Applied Animal Welfare Science, 16: 384-385.

CWU News

A window banner at Grandview School District

$4 million federal grant aimed at helping with school psychologist shortage

February 25, 2026 by

Photo of Kendall Smart working with a student at the Disability Services office

CWU Disability Services aims to provide equitable access to education

February 23, 2026 by

More News
Give to Program